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Office of Science Engagement

Tropical Field Ecology – BIOL 499A
Tropical Field Ecology – BIOL 499A
Penn State and Costa Rican faculty members will lead group research projects (four to six students per group) at sites 1 and 4. We also expect to have a marine biologist graduate student serve as the teaching assistant. Students will conceive and carry out their own research projects toward the end of our stay at Campanario. All projects are to be presented orally to the class and handed in as formal lab write-ups. Faculty will assist and interact closely with students in all aspects of the research projects.


Students are eligible to apply for:



**2020 – 2021 target application date is September 30, 2020*


 Visit the program website for more information.

Dr. Jim Marden, Professor of Biology, Penn State University

Costa Rica, Central America

Winter Break: December 27, 2020 – January 11, 2021

3 Credits

Prerequisite(s): None

Target application date: September 30, 2020

Applications will be evaluated on rolling basis until all spots are filled.

Students hiking at Volcan Arenal in Costa Rica

General Questions


124 Ritenour Building

University Park, PA 16802

Education Abroad

Vacant – Direct all questions to Tomalei Vess

Undergraduate Research

Dr. Tomalei Vess


Professional Development

Dr. Ann Marie Daniel