Office of Science Engagement

Office of Science Engagement

Tropical Field Ecology – BIOL 499A
Tropical Field Ecology – BIOL 499A
Penn State and Costa Rican faculty members will lead group research projects (four to six students per group) at sites 1 and 4. We also expect to have a marine biologist graduate student serve as the teaching assistant. Students will conceive and carry out their own research projects toward the end of our stay at Campanario. All projects are to be presented orally to the class and handed in as formal lab write-ups. Faculty will assist and interact closely with students in all aspects of the research projects.



Students are eligible to apply for:


**2018/19 application deadline is September 24, 2018*
 Visit the program website for more information.

Dr. Jim Marden, Professor of Biology, Penn State University

Costa Rica, Central America

Winter Break: December 27, 2018 – January 11, 2019

3 Credits

Prerequisite(s): None

Application deadline: September 24, 2018

Students hiking at Volcan Arenal in Costa Rica

General Questions


124 Ritenour Building

University Park, PA 16802

Education Abroad

Nimisha Thakur


Undergraduate Research

Dr. Tomalei Vess


Professional Development

Dr. Ann Marie Daniel


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